"See!" Matatu told them. "This is where Tukutela stood when he heard the indeki coming. Here and there he turned to look up in the sky and challenge us." Matatu imitated the old bull, holding his head at the same angle, humping his back and cupping his hands at each side of his head. It was such a faithful impression that for a moment he seemed to become the old bull and they all laughed. Claudia forgot her fatigue and applauded.

"Then what did the old bull do?" Sean demanded. Matatu spun and pointed along the run of the spoor.

"He went away with all his speed, he went fast and very far."

"Well," Sean said, "that puts us almost exactly forty-eight hours behind him and we have to sleep now. We'll be fifty-five hours behind him when we march again."

Tukutela's dam had been the matriarch of a herd of over one hundred beasts. She had come into her last period of estrus in her fifty-second year, and over the days it lasted, she had been mounted and serviced by six of the herd bulls, all young animals, vigorous and at the height of their powers.

It was the ideal formula for the conception of an extraordinary calf, old cow and young bull. Although it was uncertain which seed had taken root in her, the old cow had carried the genes of great elephants, big in body and tusk, natural intelligence, and the urge to dominate. These same genes had made her the leader of her herd, and now she transferred them to the fetus she carried in her womb.

She carried him twenty-two months, and then in the year when the German askaris under General von Lettow-Vorbeck were ravaging eastern Africa, the year 1915, she left the herd. Accompanied only by another old female past calf bearing, her companion of forty years, she went deep into the fastnesses of the swamps that lie on the south bank of the Zambezi River and there, on an islet fringed with ivory-nut palms, surrounded by miles of papyrus beds, and with the white-headed fish eagles chanting overhead, she cleared an area of sandy earth for her couch. When her time came, she spread her back legs and squatted over the open area, squealing in the agony of her labor, her trunk rolled up on her chest.

Her eyes had no tear ducts to drain them, so the tears poured freely down her withered cheeks as though she wept, and the spasms racked her huge, gaunt frame. The other old cow stood close beside her like a midwife, caressing her with her trunk, stroking her back and rumbling with sympathy. She forced out the calf's head and then rested for a minute before the last violent effort expelled the purple-pink fetal sac and the calf fell to the earth, rupturing the umbilical cord. Tukutela began to struggle immediately, still trapped in the glistening mucus-coated membrane, and the old cow, her companion, stood over him and, with the prehensile tip of her trunk, delicately stripped it away.

Then with her trunk his dam gently and lovingly lifted him to his feet and placed him between her front legs, making the deep, purring rumble of elephant contentment. Still wet and smooth and shining pinkly from his birthing, covered in copious gingery hair, almost blind, Tukutela rolled his little trunk back onto his forehead and reached up instinctively to the twin breasts on his mother's chest.

While he tasted the rich creamy milk for the very first time, his dam picked up the fetal sac and afterbirth and stuffed them into her mouth, chewing and swallowing and, at the same time, using her trunk to cover the damp and bloodstained spot on the earth with sand.

The three of them, his mother, her companion, and Tukutela, remained on the island for almost two weeks while the calf mastered the use of his legs and trunk, the pigment of his skin darkened, and his eyes adjusted to the harsh African sunlight. Then, when she considered him strong enough, she took him to find the herd, pushing him ahead of her and lifting him over the steep and difficult places.

The din of a hundred elephants feeding carried to them from afar, the crack and crash of breaking branches and the pig like squeals of the calves at play. Tukutela's dam trumpeted her return, and the herd came rushing up to greet her. Then, discovering the new calf, they crowded around to touch him with their trunks, puffing his scent into their mouths so they would recognize it always thereafter.

Tukutela cowered between his mother's front legs, overwhelmed by the huge bodies that surrounded him, making little baby noises of terror, but his mother draped her trunk over him and rumbled to reassure him. Within hours rather than days, he ventured out from her protection to join the other calves and to begin carving for himself a niche in the hierarchy of the breeding herd.

The herd was a close-knit group, almost all its members blood relatives, mutually reliant on each other so that the education and discipline of the young was a concern of all.

The calves were always kept in the center of the herd, and their antics were strictly supervised by the old barren cows who were their self-appointed nursemaids. Their care and protection was intense, but any infringement of the herd law was punished instantly: a tree branch wielded with gusto across the recalcitrant's back and hindquarters would ensure terrified squeals and instant obedience.

Tukutela learned his place in every situation: at the center when the herd was relaxed and feeding; between his mother's front legs when they were on the march or in flight from danger. He learned to react instantly to the alarm signal, learned to recognize it even when it was given by an animal on the further outskirts of the group.

At the signal, the instantaneous silence, in contrast to the preceding happy uproar of the herd, was an eerie phenomenon of elephant behavior.

Tukutela's development was closely parallel to the ages of a human being: His infancy lasted two years, during which time he shed the tiny milk tusks with which he had been born and entered on his juvenile years, when his true tusks emerged beyond his lips.

At first these were covered by a cap of smooth enamel, but as soon as he was weaned and began to use his tusks to feed with and to engage in mock combat with his peers, this was worn away and the true ivory beneath was exposed.

His tusks would continue to grow in length and girth throughout his entire life, even into his extreme old age. But the genes that dictated their extraordinary development came down from his dam along with all her other gifts of strength and bulk and intelligence.